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This article is about the xeno-arthropodal species. For other uses of Drone, see here.
Yanmee H2A.png
Biological overview

Latin name:

Turpis rex





Physical information

Avg. height:

177.8 centimetres (5.83 ft)–205.8 centimetres (6.75 ft)[1][2]

Avg. weight:

77.0 kilograms (169.7 lb)–127.0 kilograms (279.9 lb)[1][2]


Xeno-arthropodal morphology, beetle-like wings and exoskeleton. Flight-capable. Vestigial third pair of limbs on abdomen

Sociocultural information



Technology level:

Native Tier 4/Adopted Tier 2[3]

Notable individuals:

Also known as:

"The flying Yanme'e, simply called the Drones by some, tend to a hivelike society, eusocial and separated by castes, each with its own tasks, subject to the will of a matriarchal queen."
— From the Prophet of Clarity's Notes on the History of the Covenant[5]

The Yanme'e (Latin Turpis rex, translated to "dishonorable king"[6]), referred to by some individuals as Drones[5] or Buggers,[4] are a race of sentient, carnivorous, flight-capable insectoids.[7] The Yanme'e were one of the species that composed the religious hegemony known as the Covenant, having become the body's second client species in 1112 CE.[3] In combat, the Yanme'e served as mechanics and soldiers, often using their ability to fly to carry out ambushes on enemies of the Covenant.[1]


The Yanme'e evolved from winged tree-dwelling arthropods on Palamok which had developed complex motor skills due to a flexible appendage made up of chitin that functioned as an opposable thumb. Eventually, the species developed higher levels of technology that ultimately led to the evolution of the Yanme'e species, due to societal coordination and cooperation.[7] The Forerunners noted that the Yanme'e were one of the few sapient, arthropodal species in the Milky Way galaxy that were competent enough to attain tool-making and technological skills.[3]

The Covenant[edit]

Eventually, the Yanme'e became an interstellar-faring, Tier-4 species. The Yanme'e first encountered the Covenant in the year 1112, while the Covenant was searching for Forerunner artifacts.[8] Initially, first contact was violent as the Yanme'e attempted to resist the alien invaders. Both sides took heavy casualties, but surprisingly,[3] it was the Covenant who fared worse in these battles due to the sheer amount of Yanme'e combatants.[1] However, the Covenant was unwilling to glass Palamok, as the San'Shyuum believed that Palamok's hives could be guarding Forerunner relics on the planet's surface. The San'Shyuum eventually found an effective way to communicate with the Yanme'e, and subsequently incorporated them into the Covenant by way of a treaty in an effort to prevent a prolonged war. Exploration of Palamok following the war provided no evidence of former Forerunner artifacts on the planet.[7] The San'Shyuum were quick to make use of their talent for repairing machinery, and employed them as mechanics on Covenant vessels.[9] An assembly of Yanme'e Hive Queens was formed, and they conscripted members of the drone caste into the Covenant military.[5]

The Yanme'e were one of the first species to make contact with humanity, on the human colony of Harvest in February 2525. They were first identified by the United Nations Space Command in the siege of Gladsheim, a small town on Harvest.[10] During the later fight on the Tiara, several Yanme'e killed the Huragok Lighter Than Some out of jealousy, causing the enraged Unggoy Deacon Dadab to slaughter them in revenge.[11]

After the Hierarchs declared war on humanity, Yanme'e were used as military combatants against UNSC forces. They were deployed during many major conflicts, including the Fall of Reach. During the Battle of Meridian, five or six Yanme'e were deployed by an SDV-class heavy corvette to finish off the survivors of a boat it had fired upon. Dorian Nguyen was able to kill one by slamming it with a guitar, crumpling its wings and sending the Yanme'e into the sea. Dorian managed to escape, but the Yanme'e slaughtered the hundreds of other people on the boat, including Dorian's friends and bandmates.[12]

During the Battle for Earth, they boarded orbital defense platforms along with other Covenant infantry.[13] When the Covenant made landfall in New Mombasa, the Yanme'e immediately established a large hive in the Superintendent's data center.[14]

During the Great Schism, the Yanme'e joined the San'Shyuum and the Jiralhanae.[15] Many were present as combatants during the Battle of Installation 00.[16]


After the Human-Covenant War, many Yanme'e happily returned to their home system.[7] In 2554,[17] Many Yanme'e served in Tem'Bhetek's faction which attempted to use the Prototype Halo installation to destroy Sanghelios and the Sangheili. Many were killed by Rtas 'Vadum and Tul 'Juran after mortally wounding Tem'Bhetek with the remained trying desperately to get to Boru'a'Neem to stop him from firing the miniature Halo again. They were killed when Tem destroyed the installation.[18] A large number of Yanme'e were present during the Battle of Draetheus V, serving in Merg Vol's Covenant.[19] Many more served in Jul 'Mdama's Covenant during the Battle of Installation 03.[20] For the most part, though, the Yanme'e mostly kept to themselves. The UNSC attempted a "don't poke the nest" policy toward the Yanme'e, attempting to avoid direct provocation while the UNSC focused on more immediate threats within the Covenant remnants. This policy was largely successful, though there were exceptions.[21]

When Cortana made her offer at the beginning of the Created conflict to the species of the galaxy, she did not include the Yanme'e in her list of species that she offered to fall under her protection. Upon hearing her message, Spartan Edward Buck noted her exclusion of the Yanme'e and didn't blame Cortana for excluding them as the Yanme'e creeped him out.[22][23]


Anatomy and physiology[edit]

Diversity of Yanme'e protectors hanging asleep from a Phantom.
"They're built kinda like beetles, and they flit around on twinned pairs of transparent wings that flap faster than you can see."
Edward Buck on the Yanme'e[24]

The Yanme'e share many similarities with Earthly beetles and colonial insects. They are bipedal arthropods with hard exoskeletons made from a glossy chitinous substance, as well as two pairs of veined wings. These adaptations are a response to the high gravity of the Yanme'e homeworld, Palamok; their shells allow them to withstand hard falls, and the thick atmosphere of Palamok allows the winged Yanme'e to achieve just enough lift to perform fluttering leaps or naturally short bursts of flight.[25] In lower-gravity environments, Yanme'e are capable of using their wings for actual aerial maneuvering, but the weight of their exoskeletons prevent them from flying.[1] When the Covenant discovered this natural ability, scientists extended their flight capabilities by fixing gravity-dampening modules to their bodies. The module adjustment gave the Yanme'e the ability to take flight freely, and the species proved to be impressive fliers—particularly on worlds that had less gravity than Palamok.[7] Yanme'e are also impressively strong, despite their frail appearance. They can also be seen lifting their enemies off the ground to impressive heights.[24][26]

Unlike Earth insects, which have three body segments, Yanme'e have five: a head, a cephalothorax, a thorax, a pelvis, and an abdomen. The head supports a pair of feathery sensory antennae and luminous,[27] compound eyes that give the species adept vision.[28] The mouth consists of several sets of mandibles like those found on insects. Their hands sport opposable thumbs and three dextrous fingers. Yanme'e are "polydextrous", meaning that they are capable of using both their prehensile hands and feet to operate tools. The primary pair of legs are attached to a pelvis-like structure, while a smaller pair is attached to the abdomen. Although they generally use their primary set of limbs, Yanme'e can use this last set of limbs to operate weapons, grasp enemies, or latch on to ceilings and hang down vertically during sleep.[7][29] The Yanme'e have extremely sharp claws, located on all limbs,[30] that they can use to defend themselves if necessary, though they prefer to avoid close-quarters combat.[31] Yanme'e are a carnivorous species. Early in their history, the Yanme'e developed and bred types of feral livestock for food.[3]

There is essentially no apparent sexual dimorphism between male and female Yanme'e. Yanme'e exoskeletons are brightly colored, with different hues representing different functions or class.[7] Adolescent, or pupal, Yanme'e have translucent shells that glow luminescent yellow and orange. Almost nothing is known about the Yanme'e life cycle beyond the fact that the queen lays eggs. It is possible that the Yanme'e go through a complete metamorphosis including an egg, larva, pupae and adult stage, as many terrestrial insects and arthropods do.


Yanme'e are a eusocial species, and therefore their intelligence is geared toward working in concert with large groups. They can accomplish tasks swiftly and efficiently when working together, and their knowledge of machinery is rivaled only by the Huragok. However, they lack individuality because of their hive lifestyle; their instincts and societal conditioning suppress creative thought and encourage total obedience to authority. They do not appear to experience higher emotions; it is known that they are able to feel jealousy, as seen when they killed the Huragok Lighter Than Some because he usurped their role as technician on board Rapid Conversion.[32] Yanme'e were allegedly incapable of feeling fear.[33] Despite their hive-minded nature, Yanme'e are still individually quite intelligent.[24]

Certain Yanme'e are unable to function in a hive society due to a form of personality disorder; these "Unmutuals" are capable of independent thought and tactical planning, although they tend to be extremely violent and aggressive. However, some of these Yanme'e appear to be very intelligent independently.[34] One individual was able to tamper with the inner circuitry of a UNSC Interrogator and allowed the device to translate the Yanme'e language into English in a matter of seconds.[35]


The Yanme'e are an insectoid species with a hive-minded social structure. Their society consists of reproductive queens, reproductive male drones and non-reproductive workers and soldiers. It is unknown if the soldier and worker caste is specific to one gender or another, but it is confirmed that the Yanme'e worker Hopalong was a male. Because of their eusocial nature, they can be extremely dogmatic and hive-minded,[28] a trait that may have made for an easy conversion into the Covenant hierarchy. The Hive Queen is the center of Yanme'e society and culture, to the point of her being the reason for the species' technological progress and advancement. This includes most major advances in science and other technology that typically results in emphasized protection and augmentation of the queen and the hives.[7]

Yanme'e obeyed the commands of the higher castes without question in the Covenant, but had little concept of religion and did not partake in social norms due to communication difficulties with other species. They referred to the San'Shyuum as "Queens", a remnant of their former hive lifestyle.[3] They did not interact with other Covenant member races except to trade resources and to serve in the military, where they typically serve as mechanics and repair crews on warships. Although highly skilled with technology, they aren't nearly as effective as Huragok; this has often brought them into a state of rivalry with Huragok, and in jealousy, the Yanme'e aboard Rapid Conversion even killed a Huragok named Lighter Than Some when it was assigned all of their former tasks.[32]

Role within the Covenant[edit]

A Yanme'e combat swarm on Reach.

Yanme'e were initially conscripted into the Covenant only as mechanics and laborers. The Yanme'e were not as technically skilled as the Huragok, but they still proved to be incredibly competent mechanics and were efficient at general maintenance.[36] After years of serving in only custodial positions within the Covenant military, the Covenant had eventually developed numerous combat roles for the Yanme'e—largely due to their impressive flight capabilities.[7] They are typically used for surprise aerial ambushes and assaults, such as catching infantrymen off guard and diverting their attention so that their Covenant allies may gain the upper hand. The Yanme'e physiology made the species well-suited in infiltration roles.[33] Yanme'e are quite capable of using most light Covenant weaponry, and depending on their rank may wield plasma pistols, Needlers, and plasma rifles. The limb dexterity of the Yanme'e allows them to have competent aiming skills while in flight or while they are climbing over a structure. In addition, their speed and adept sight made the Yanme'e one of the most threatening infantry units in the entire Covenant military.[3]

Yanme'e were very territorial over the tasks they were assigned to. Although the Yanme'e would respect the Covenant's religious principles and would loyally obey their Sangheili and San'Shyuum commanders, they had no particular passion for the Covenant religion or desire to follow Covenant social normalities. Due to their lack of individual initiative, the Yanme'e remained low in the Covenant hierarchy, often commanded on the battlefield by Sangheili, Jiralhanae, or Kig-Yar.[31] Due to their low position, the Yanme'e were not represented on the High Council, but their interests were instead vetted by the Ministry of Concert.[37]

During large-scale ground engagements, Yanme'e have a tendency to entrench themselves and establish hives within enemy territory, as seen during the Fall of Reach and the conflicts in New Mombasa. This can make them a persistent threat, as the hives provide a constant supply of Yanme'e to reinforce occupying Covenant troops.[14]


A Yanme'e hive under New Mombasa.
"The first thing the Drones usually do when they set up in a place is build a hive made of spit, shit, and whatever else they can find lying around. It's how they mark a conquered area as their own."
— Edward Buck[24]

Yanme'e have the ability to form fully functional hives at incredibly fast rates; during the Battle of New Mombasa, the Yanme'e generated a vast hive structure within the city's data center within hours. Hives are primarily constructed from a collection of mud, wood, and cellulose materials that is then mixed with Yanme'e excretions, which are formed into large stalactite-shaped structures. Yanme'e are able to establish hives in any location where sufficient raw material was available. However, hives are most commonly found underground or in vast jungle canopies, like the ones that cover a majority of Palamok's surface. The Yanme'e have also formed densely packed hives within the massive geological structures on Palamok's moons of Ka'amoti and Oquiu.[7] A defining characteristic of Yanme'e hives is that they radiate heat to their surroundings from vent-like openings, elevating the temperature over a large area.[14]

During their service with in the Covenant, Yanme'e that are deployed in combat roles are often ordered to create temporary, makeshift hives that allowed the rest of the Covenant military to easily access enemy territory. The hives served as Covenant strongholds for invading forces, often hidden within enemy territory on cavernous ceilings or underground.[7] Hives were established on Earth and Verge by the Yanme'e during the Human-Covenant War.[14][38] They would often use these hives to show that they had conquered an area on a world.[24]


The Yanme'e language is a cacophony of high-pitched clicking or whining from the rubbing of their waxy, leaf-like wings.[39] They are also known to communicate sonically with loud shrieks and screeches, as well as through the use of pheromones.[1][7] Combat units will usually elect an attaché to serve as a translator, learning how the other races communicate and relaying the relevant information to the other Yanme'e. However, the attaché still does not socialize with other species, only learning enough about their allies to keep the other Yanme'e organized and informed.[31] By 2552, the Yanme'e language was still not completely translated by the Office of Naval Intelligence due to lack of information about the species. ONI's Interrogator translation software was only capable of poorly imitating Yanme'e clicks, but the output was unintelligent to the species.[35]


The Yanme'e race, inclining towards a hive-like and eusocial society, are divided by several castes. Most individuals fall into one of the three main, distinct Yanme'e castes: domestic, worker, or protector. The colorations of the Yanme'e shell will indicate its caste, and function within the caste. All castes are subservient to the Hive Queen.[5][7]

Hive Queen[edit]

Main article: Yanme'e Queen

The Hive Queen is the central figure of the hive. Her primary role, as with ants and bees, is to propagate the species. The queen serves as the leader of the Yanme'e species. The Yanme'e lead an aggressively matriarchal society, with the other castes of the Yanme'e serving the queen and carrying out her orders to ensure the preservation of the hive. Yanme'e queens are responsible for the species reproduction and way of life, leading to the well-being of a queen being the uppermost importance of a hive.[5][7] The queen substantially larger than the other castes of the race, with the distended, egg-producing abdomen and thick legs.[40] It is unlikely that a single queen could support an entire species so there are probably multiple, though possibly subordinate, queens.

Protector caste[edit]

Yanme'e soldiers from the protector caste.

Similar in function to the soldier caste within many Earth-based insect species, the protector caste is charged with defending the queen. This caste was artificially merged with the worker caste by the Covenant, following the Yanme'e's joining of the Covenant.[7] The protector caste was used as the Yanme'e military, within hive society and in the Covenant military. Within the protector caste, there appear to be many ranks. Minors are the most common form and have a very low position in Covenant society. Majors likely represent higher class in the protector caste: they have larger wings, fly faster, are more skilled in combat and often carry Needlers.[41]

Worker caste[edit]

The worker caste is similar to the worker castes of similar function found within many eusocial Earth-based insect species. This caste was artificially merged with the protector caste by the Covenant, following the species' incorporation into the Covenant.[7] The caste is likely responsible for creating the hives and were likely employed by the Covenant in custodial positions. When the Covenant forced Yanme'e workers into combat positions on Rapid Conversion, several individuals became unusually clumsy and flustered; they became confused over the change in task.[28]

Other castes[edit]

Yanme'e males that are unable to fit in with the caste system and are born with destructive tendencies are known as Unmutuals and are exiled from hives.[7] These are Drones unable to work in concert with the rest of the hive, and have in fact been known to go on rampages destroying nests, smashing eggs, even murdering other adult Yanme'e. They are generally imprisoned to prevent them from communicating with the rest of the hive, and on at least one known occasion have been sent to mine helium-3 on planets captured by the Covenant. Yanme'e in these penal colonies are often guarded by Kig-Yar who may work them to death. On one occasion a Kig-Yar actually removed the legs of a drone whom SPARTAN Team Black named Hopalong. He helped the SPARTANs infiltrate their objective, deceiving them long enough to release the rest of the drone prisoners.[34]

The domestic caste forms another large division within the Yanme'e caste structure, but its functions remain unknown. It is likely related to the wingless males that serve the queens.[40]

Homeworld and colonies[edit]

Main article: Palamok

The Yanme'e homeworld is Palamok, an Earth-like planet but twice the size with over twice the gravity (2.2 G). Palamok is a flat and geographically featureless world, with little tectonic or volcanic activity.[3] The planet features a warm atmosphere and a stable environment.[25] The planet's high gravity led to the Yanme'e evolving with hard shells that are able to absorb a large amount of damage. Due to their planet's high gravity, the Yanme'e are only capable of short bursts of flight on Palamok, though they can fly quite well in 1.0 G environments.[2] The Yanme'e have several colonies throughout the Napret system, with Oquiu and Ka'amoti being the most prominent.[7]

Military structure[edit]

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Though a sense of organization or leadership is usually unclear among swarms of attacking Yanme'e, rank structure can be surmised based on the color of a Drone's exoskeleton and the strength of its defensive armor. The reason for the association of color with strength and rank is unknown, but one possible theory is that it is based on age. Yanme'e, like many real-world insects, may grow by molting their skins. The color changes could occur after each molt, when the Drone becomes larger and stronger.

An attaché is selected from each Yanme'e combat unit and is tasked with interfacing with the other Covenant races in order to organize the swarm. Due to the difficulty in communication among the Yanme'e and the other Covenant races, these attachés do not socialize with their superiors. Instead, they are expected to become familiar enough with their allies' modes of communication that they can keep their unit mates coordinated effectively.[1]

Rank Image Description
Yanme'e Leaders
Yanme'e Queen Queens are the leaders of Hives, responsible for the birth of new members of the Swarm.
Yanme'e Leader Drone Leader.jpg Drone Leaders appear to command groups of Yanme'e Minor and Majors.
Yanme'e Infantry
Yanme'e Ultra Yanme'e Specialist (blue - reach) 2.jpg An Ultra is considered the highest regular military rank for the species. They are distinguished by their blue and cyan exoskeletons, along with paired forked horn-like structures on their head and cephalothorax.
Yanme'e Major ReachRedShell.jpg Drone Majors are one step higher than the Minor Drones. They still appear low in rank, about the same as a Jackal Minor. They are most often seen commanded by Jackal Majors, Brutes, and even Special Operations Grunts. Yanme'e Majors are differentiated from lower ranks by their crimson shells and energy shields. They have longer wings and can fly faster than Drone Minors.
Yanme'e Minor Yanme'e Minor (green) 2.jpg Drone Minors are the lowest of the Drone ranks. They are commanded by Drone Majors, Jackals, and Grunt Majors. They seem equal, if not even inferior in rank, to the Grunt Minors, placing them at the very bottom of the Covenant hierarchy. Usually found equipped with Plasma Pistols, Needlers, and rarely Plasma Rifles, Drone Minors, while powerful in swarms, are weak individually. They can be taken down fairly easily, and will often make tactical errors such as flying into an enemy's line of sight, or getting in the path of another Drone's shot. They have green, blue, and silver exoskeletons.

Other Ranks[edit]

Regular Drone infantry are composed of the following ranks:

  • Green - Green Drones are the most common type of Yanme'e encountered and are by the far the weakest.
  • Blue - Blue Drones are moderately common on the battlefield but are slightly stronger than their green counterparts.
  • Silver - Silver Drones are both slightly more rare and more resistant to attack than blue Yanme'e.

Among these infantry ranks, an additional group of leader Drones have also been seen on the battlefield, utilizing Plasma Rifles and low-powered personal energy shields to make them more capable in combat. Only two of these leader varieties have been seen:

  • Gold - Gold Drones are adolescent leaders who have been rushed from their pupal stage. As a result, their abdomens glow brightly.[42] They occur in slightly less numbers than their red brethren.
  • Red - Red Drones represent the peak of the Yanme'e ranks. They are extremely capable warriors compared to the rest of their brothers and feature weak energy shields.


Language Equivalent
Flag of France.png

Les Drones in all media.

Les Vermines (nickname) in Halo: Reach's manual.

  • The nickname "buggers" is likely a reference to the Formic race in the Orson Scott Card novel Ender's Game. Additionally, the Drones' native name, Yanme'e, could be based on the Japanese 蜻蜓 (yanma), meaning "large dragonfly".
  • The weight of a Drone averages around 77 kilograms, a reference of Bungie's love of the number 7.
  • During concept stages for Halo 3, the Yanme'e were planned to have an animation in which they would carry away a Marine.[43] This animation was cut from the final game but one does pick up a Marine for a short time at the beginning of the first swarm attack in Crow's Nest.
  • In Halo: Reach, when Drones are killed with a headshot, they explode, leaving behind only limbs.
  • In Halo: Reach, Drones are not encountered in Firefight mode, unlike Halo 3: ODST.
  • On the game stats for Halo: Reach, Yanme'e are referred to as "Buggers" rather than their more common nickname, "Drones".


List of appearances[edit]


  1. ^ a b c d e f g Halo Encyclopedia, pages 148-149
  2. ^ a b c Halo: The Essential Visual Guide, page 51
  3. ^ a b c d e f g h Halo 3, Bestiarum
  4. ^ a b c d Halo 2, Conversations from the Universe
  5. ^ a b c d e Halo: Broken Circle, page 186
  6. ^ Bestiarum Latin names translated
  7. ^ a b c d e f g h i j k l m n o p q r Halo Waypoint: Yanme'e
  8. ^ Halo Encyclopedia, page 159 (2011 edition)
  9. ^ Halo Encyclopedia, page 31
  10. ^ Halo: Contact Harvest, page 286
  11. ^ Halo: Contact Harvest
  12. ^ Halo: Battle Born, Chapter 4
  13. ^ Halo 2, campaign level, Cairo Station
  14. ^ a b c d Halo 3: ODST, campaign level, Data Hive
  15. ^ Halo 2, campaign level, Uprising
  16. ^ Halo 3, campaign level, The Ark
  17. ^ Halo Waypoint: Catalog Interaction (Page 43)
  18. ^ Halo: Shadow of Intent
  19. ^ Halo: Spartan Assault
  20. ^ Halo: Spartan Strike
  21. ^ Halo: New Blood, Chapter 16
  22. ^ Halo 5: Guardians, campaign level Guardians
  23. ^ Halo: Bad Blood, page 5
  24. ^ a b c d e Halo: New Blood, pages 69-70 (Google Play edition)
  25. ^ a b Halo Encyclopedia, page 309 (2011 edition)
  26. ^ Halo 3, campaign level, Crow's Nest
  27. ^ Halo: Contact Harvest, page 202
  28. ^ a b c Halo: Contact Harvest, page 267
  29. ^ Halo 3: ODST
  30. ^ Halo: Contact Harvest, page 371
  31. ^ a b c Halo Encyclopedia, page 159
  32. ^ a b Halo: Contact Harvest, page 369
  33. ^ a b Stats
  34. ^ a b Halo: Evolutions, "Blunt Instruments", page 26
  35. ^ a b Halo: Evolutions, "Blunt Instruments", page 13
  36. ^ Halo Encyclopedia, page 121
  37. ^ Halo: Contact Harvest, page 150
  38. ^ Halo: Evolutions, "Blunt Instruments", page 21
  39. ^ Halo: Contact Harvest, page 311
  40. ^ a b Halo: Contact Harvest, page 381
  41. ^ Halo: Reach
  42. ^ Halo 3: ODST Official Strategy Guide
  43. ^ A Certain Sense of Movement