Rapid Conversion was originally decommissioned by the Sangheili for their own use, but was deemed worthy of use by Jiralhanae. Many of its systems, including weapons and gravity lifts, were disabled. Upon taking command of the vessel, Chieftain Maccabeus commissioned a number of changes to its interior decor, including an authentic facsimile of a traditional Jiralhanae feasting hall with doors and ceiling supports of forged steel and lighting provided by primitive oil lamps on clawed floor stands. The crew was comprised of a pack of ten Jiralhanae, roughly sixty Unggoy, and over a hundred Yanme'e responsible for the vessel's upkeep.
The warship was used to deploy troops and vehicles to the surface of Harvest during the First Battle of Harvest and also used as the Covenant's command center. Though its weapon system, a plasma torpedo turret, had been disabled, the Huragok Lighter Than Some was able to repair them in time for the ship's actions against Harvest and its colonists.
Later, going into low orbit over Harvest's surface in order to deliver troops and gather the relics they sought, Rapid Conversion fell into a trap that had been laid by Captain Ponder and his troops and was hit by two rounds from a mass driver based on Harvest's surface. The first round did little damage. The second round, however, passed completely through the warship, severing the connections between the ship's reactor and anti-grav generators, causing the ship to plummet to the planet's surface.
In the following months however, the crew managed to repair the ship and began to glass Harvest, eventually reducing the planet's surface to cinder. In April, Rapid Conversion destroyed the Colonial Military Administration scout ship Argo. Six months later, the CMA's Battle Group 4, consisting of the Heracles, the Arabia, and the Vostok, was sent to investigate the loss of contact with Harvest and the Argo; the battle group was almost completely destroyed by a Covenant super-destroyer that was holding station over the planet.
After ascending to the title of Chieftain of the Jiralhanae, Rapid Conversion remained under Tartarus' control. During the Great Schism, following Tartarus' death on November 3, 2552, Sangheili assassins led by Resa 'Azavayl under the banner of the Silent Shadow eliminated the remaining members of Tartarus' crew aboard the ship.
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