From Halopedia, the Halo wiki
Rangmejo (as they were known to Forerunner Lifeworkers), or "quadwings", are a species of large pterosaur-like creatures with dual sets of wings. They are among the most ubiquitous varieties of fauna that have been encountered on Forerunner installations by the United Nations Space Command, as they have been observed on Installation 00 and multiple Halo rings. Rangmejo have been known to gather in significant numbers within artificially-constructed chambers.
Rangmejo were among the creatures preserved by the Librarian against the firing of the Halo Array in 97,445 BCE. Following this destructive event they were seeded upon at least two Halo rings, installations 03 and 05, while also being left to flourish on the Ark, Installation 00, far beyond the Milky Way galaxy. Rangmejo had been specifically selected and adapted by Lifeworkers to play diverse and important roles within a wide range of ecosystems, helping to contribute to the overall health and stability of various biomes.
In November of 2552 rangmejo were observed by the battling forces of the Covenant and United Nations Space Command when their war brought them to Installation 05. In December of that same year an unfinished Halo ring was fired above the Ark, spelling death for the many sentient creatures which inhabited its surface. The Ark's monitor, 000 Tragic Solitude, eventually then began to breed new life forms. Among these were more rangmejo. Following the discovery and subsequent exploration of Installation 03 by humanity in 2552 and 2553, rangmejo were witnessed there. Respected scientist Doctor Luther Mann's observations of those on Gamma Halo made him certain it would not be safe to get close to them.
In March of 2555 a small group of humans and Sangheili representing the United Nations Space Command and Swords of Sanghelios, respectively, encountered a large number of the creatures within a dimly lit cavern near the Ark's Foundry. Fortunately for them, the rangmejo left abruptly soon after they were sighted.[note 1] By 2558 the UNSC had procured a few individual specimens for scientific study.
Anatomy, physiology, and behavior
Rangmejo are large creatures with two sets of impressive wings flanking a narrow, muscular body. At one end of that body is a head featuring a strong, lengthy beak and a protruding crest—often brightly colored. At the other is a long tail that can represent over half of the length of the individual's body. Their considerable size is matched by their two large wingspans, the wings of which flap and beat heavily. Rangmejo somewhat resemble Pteranodon species, thanks to their prominent head crests. Rangmejo blood is red. The specimens acquired by the UNSC show remarkable evidence of genetic optimization, their ancestors having been specially adapted by Forerunner Lifeworkers. Rangmejo have been noted to make a squealing sound while flocking together in large numbers.
Behind the scenes
Quadwings first appeared in Halo 2, specifically within the campaign missions Delta Halo and Regret. They are absent from Halo 3's campaign but are featured in two of its multiplayer maps: Epitaph and Guardian. Within these games it is possible for players to kill the rangmejo flying about, resulting in a bloody eruptions. When a Halo-themed texture pack was released for Minecraft in 2014, it allowed the ender dragon in that game to be re-skinned to resemble a quadwing. Quadwings were first identified as rangmejo the following year within a Canon Fodder entry which followed the release of Halo: Hunters in the Dark. 343 Industries provided a digital asset to Falcon's Creative Group so that it could feature quadwings in "The Ring Experience" at Halo: Outpost Discovery, but the creature ultimately had to be recreated due to what already existed being too undetailed and unrealistic.
List of appearances